This article first appeared in the by Jennifer Mueller,
M.S. Candidate The next time you browse through the
produce department in your local supermarket, take a closer look at some of
those fruits and vegetables. They contain substances called bioflavonoids,
which are part of a larger group of molecules called polyphenols. These
flavonoids play a major role in the color and taste of many of the fruits and
vegetables we eat, and they possess numerous medical benefits as well. History Within the last few decades, extensive
studies have been conducted on a wide range of flavonoids, finding a variety
of beneficial effects. Anthocyanidins (found in red wine and pomegranates)
have demonstrated anti-cancer, anti-inflammatory, and anti-microbial effects
.2,3 Gallates and catechins (found in green teas) have shown anti-cancer
effects as well.4 Quercetin, rutin and others have shown strong protective
effects against lipid peroxidation,2 and quercetin has also been proven
effective in the prevention and treatment of diabetes-induced cataracts.5
Silymarin (found in milk thistle) is used in Europe as a prescription drug to
treat various liver diseases, and cyanadin and catechins have also shown
similar hepatoprotective effects.2,3 Numerous flavonoids have demonstrated
potent anti-microbial and anti-viral effects, and some, such as quercetin and
morin, have shown to be effective in lowering cholesterol in the blood.6
While numerous beneficial effects of flavonoids have been examined, perhaps
one of the best documented medical applications is their anti-allergic
effects. Allergies and Flavonoids Bioflavonoids
have long been known to possess anti-allergic effects. As early as the
1950's, studies showed that flavonoids could prevent the release of
histamines and inhibit anaphylaxis.8 It wasn't until some years later,
however, that any more attention was paid to the link between flavonoids and
allergy. The breakthrough was the discovery of the flavonoid khellin that
turned out to be an effective muscle relaxant. During attempts to improve its
muscle-relaxing capabilities, a series of derivatives was synthesized. One of
these derivatives was disodium cromoglycate, Cromolyn. Studies suggest that
Cromolyn could alleviate the symptoms of asthma and reduce the need for other
asthma medications when taken over long periods of time. It acts by
decreasing the tran-sport of calcium into the cells, thus slowing the release
of histamine.10 Preliminary studies suggest that Cromolyn is effective in
reducing the symptoms of allergy, both when taken via inhalation and when
applied topically in the eyes.10 Other derivatives of khellin have also
been shown to be effective against asthma and allergy. The experimental
bioflavonoid-derivative drugs FPL 52757 and FPL 57787 were given orally to
rats who were then exposed to an allergen. Both substances showed protection
(similar to that seen with Cromolyn) against symptoms to the
chemically-induced allergic reaction.11 FPL 55712 given via aerosol reduced
asthmatic symptoms in guinea pigs,12 and FPL 52694 given orally to rats
inhibited reactions to allergens injected under the skin. The mode of action
for all these experimental bioflavonoid derivatives appears similar to
Cromolyninhibition of histamine release.13 Natural vs. Synthetic In vitro studies (done in cell
cul-tures instead of whole animals) have been conducted to determine the
exact mode of action by which flavonoids suppress the symptoms of allergy and
asthma. As mentioned earlier, the symptoms are mediated by the release of
histamine, leukotrienes, and prostaglandins from the immune cells. In one
study, mast cells were treated with eleven different flavonoids, including
quercetin and catechins. Histamine release was chemically induced. All the
flavonoids tested exhibited some level of protective effect against the
release of hist-amine by pre-venting the uptake of calcium into the cells.15
Similar results have been shown in basophils treated with quercetin, which is
chemically similar to the synthetically derived Cromolyn. Quercetin was quite
effective at inhibiting the release of histamine from these cells, even at
low doses.16 It is interesting to note that in these and other studies,
quercetin has been proven more effective at blocking histamine release than
the synthetically derived Cromolyn.4 While the exact mechanism by which
flavonoids prevent calcium influx into the cells is still debated, it is
thought that flavonoids may inhibit the enzyme cyclic AMP phosphodiesterase.
This results in raised levels of cyclic AMP inside the mast cells, preventing
the release of the histamine.17,18 Other Benefits As can be seen, bioflavonoids
are highly effective anti-allergy and anti-asthma agents. While more research
is needed to determine exact doses, the best methods of delivery (oral versus
inhaled), and which combinations of flavonoids can offer the best effects, it
is clear that bioflavonoids offer allergy sufferers a new form of relief,
without the annoying side effects of prescription drugs. When taking into
account all the other ways they can actas anti-inflammatories, anti-viral and
anti-microbial agents, and even anti-cancer agentsit is clear that
bioflavonoids offer a wide range of possibilities for the future of medicine. Jennifer Mueller is a Masters
Candidate in Nutrition Science at the University of California at Davis. Her
primary focus is developmental nutrition and prenatal health. She was
Managing Editor for the Journal of Optimal Nutrition and has studied under
Dr. Brian Liebovitz, the noted medical nutritionist. References: 1. Szent-Gyorgyi A, Rusznyak S. Vitamin P: Flavonoids as
vitamins. Nature 138: 27, 1936. 2. Cody V, Middleton E Jr, Harbone J (editors). Plant
Flavonoids in Biology and Medicine, vol 1. Alar R. Liss, Inc. New York, NY
1986. 3. Cody V, Middleton E Jr, Harbone J, Beretz A (editors).
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York, NY 1988. 4. Liebovitz B, Mueller J. Bioflavonoids and polyphenols:
medical applications. J Opt Nutr 2(1): 17-35, 1993. 5. Varma S, Kinoshita J. Inhibition of lens aldose reductase
by flavonoids: their possible role in the prvention of diabetic cataracts.
Biochem Pharmacol 25: 2505-13, 1976. 6. Yugarani T, Tan B, Teh M et al. Effects of polyphenolic
natural products on the lipid profiles of rats fed high fat diets. Lipids
27: 181-86, 1992. 7. Schoenkerman B, Justice R. Treatment of allergic disease
with a combination of antihistamine and flavonoid. Annals of Allergy 10:
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histamine toxicity, anaphylactic shock and histamine-enhanced capilary
permeability to dye. J Allergy 21: 133-47, 1950. 9. Davies B, Moodley I. Antiallergic compounds. Pharmacol
Therap 17: 279-97, 1982. 10. Bernstein I, Johnson C, Tse C. Therapy with Cromlyn
sodium. Annals Internal Med 89: 228-233, 1978. 11. Augstein J, et al. New orally effective chromone
derivatives for the threatment of asthma. Agents and Actions 7(4): 443-45,
1971. 12. OĠDonnell M, Welton A. Pharmacologic properties of FPL
55712 administered by aerosol. Agents and Actions 14(1): 43-48, 1984. 13. Wells E, et al. The anti-allergic effects of FPL 52694.
Int Arch Allergy Appl Immun 76: 188-90, 1985. 14. Nagai H, Osuga K, Koda A. Inhibition of hypersensitivity
reactions by soluble derivatives of baicelin. Japan J Pharmacol 25: 763-72,
1945. 15. Amallal M et al. Inhibition of mast cell histamine
released by flavonoids and biflavonoids. Planta Medica 16-20, 1985. 16. Middleton E Jr, Drzeqiecki G, Krishnarao D. Quercetin: an inhibitor
of antigen-induced human basophil histamine release. J Immunol 127(2):
546-50, 1981. 17. Beretz A, Anton R, Stoclet J. Flavonoid compounds are
potent inhibiters of cyclic AMP phosphodiesterase. Experentia 34: 1054-55,
1978. 18. Middleton E Jr, Kandaswami C. Effects of flavonoids on
immune and inflammatory cell functions. Biochem Pharmacol 43: 1167-79,
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