1520s, "moldy, sour," perhaps a variant of moisty "moist, damp" (see moist), but musty, of bread, "containing must" is attested mid-15c., and from 15c.-19c. must also was a variant of musk. Related: Mustiness.
a moldy (or musty) odor
mustee
mustelid
musteline
muster
mustn't
musty
mutability
mutable
mutagen
mutant
mutate