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HYDROCHARIDEAE

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Originally appearing in Volume V14, Page 113 of the 1911 Encyclopedia Britannica.
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HYDROCHARIDEAE , in See also:

botany, a natural See also:order of Mono- cotyledons, belonging to the See also:series Helobieae. They are See also:water- See also:plants, represented in See also:Britain by See also:frog-See also:bit (Hydrocharis Morsus- ranae) and water-soldier (Stratiotes alaides). The order contains about fifty See also:species in fifteen genera, twelve of which occur in fresh water while three are marine: and includes both floating and submerged forms. Hydrocharis floats on the See also:surface of still water, andhas rosettes of See also:kidney-shaped leaves, from among which See also:spring the See also:flower-stalks; stolons bearing new See also:leaf- rosettes are sent out on all sides, the plant thus propagating itself in the same way as the See also:strawberry. Straliotes abides has a rosette of stiff See also:sword- like leaves, which when the plant is in flower project above the surface; it is , also stoloniferous, the See also:young rosettes sinking to. the bottom at the beginning of See also:winter and rising again to the surface in the spring. Vallisneria (See also:eel-grass) contains two species, one native of tropical ,See also:Asia, the other . in- habiting the warmer parts of both hemi- See also:spheres and reaching as far See also:north as See also:south Frog-bit—male plant. the mud at the bottom I, See also:Female flower. of fresh water, and the 2, Stamens, enlarged. See also:short See also:stem bears a 3, Barren See also:pistil of male flower, enlarged. cluster of See also:long, 4, Pistil of female flower. g, narrow 5, See also:Fruit. grass-like. leaves; new 6, Fruit cut transversely. plants are formed at 7, mod' the end of See also:horizontal 8, 9, Floral diagrams of male and female See also:flowers respectively. runners. Another type s, Rudimentary stamens. is represented by Elodea canadensis or water-See also:thyme,which has been introduced into the See also:British Isles from North See also:America. It is a small, submerged plant with long, slender branching stems bearing whorls of narrow toothed leaves; the flowers appear at the surface when mature. Halophila, Enhalus and Thalassic are submerged maritime plants found on tropical coasts, mainly in the See also:Indian and Pacific oceans; Halophila has an elongated stem rooting,at the nodes; Enhalus a short, thick rhizome, clothed with See also:black threads resembling See also:horse-See also:hair, the persistent hard-bast strands of the leaves; Thalassic has a creeping rooting stem with upright branches bearing crowded strap-shaped leaves in two rows. The flowers spring from, or are enclosed in, a spathe, and are unisexual and See also:regular, with generally a calyx and corolla, each of three members; the stamens are in whorls of three, the inner whorls are often barren; the two to fifteen carpels See also:form an inferior ovary containing generally numerous ovules on often large, produced, parietal placentas. The fruit is leathery or fleshy, opening irregularly. The seeds contain a large embryo and no endosperm.

In Hydrocharis (fig. I), which is dioecious, the flowers are See also:

borne above the surface of the water, have conspicuous See also:white petals, contain See also:honey and are pollinated by in-sects. Stralioles has similar flowers which come above the surface only for See also:pollination, becoming sub-merged again during ripening of the fruit. In Vallisneria (fig. 2), which is also dioecious, the small male flowers are borne in large See also:numbers in short-stalked spathes; the petals are See also:minute and See also:scale-like, and only two of the three stamens are !fer- FIG. z.—Vallisneria spiralis—Eel grass--See also:tile; the flowers about i natural See also:size. A, Female plant; B, become detached Male plant. before opening and rise to the surface, where the sepals expand and form a See also:float bearing the two projecting semi-erect stamens. The female flowers are solitary and are raised to the surface on. a long, See also:spiral See also:stall:; the ovary bears three broad styles, on, which some of the large, sticl.•y See also:pollen-grains from the Iloatmg male floc: ers get de-posited (fig. 3). After pollination the female flower becomes See also:drawn below the surface by the spiral See also:con-See also:traction of the long stalk, and the fruit ripens near the bottom. Elodea has poly- FIG. 3.

gamo.us flowers (that is, male, female and hermaphrodite), solitary, in slender, tubular spathes; the male flowers become detached and rise to the surface; the See also:

females are raised to the surface when mature, and receive the floating pollen from the male. The flowers of Halophila are submerged and apetalous. The order is a widely distributed one; the marine forms are tropical or subtropical, but the fresh-water genera occur also in the temperate zones.

End of Article: HYDROCHARIDEAE

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