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PRIMULACEAE

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Originally appearing in Volume V22, Page 342 of the 1911 Encyclopedia Britannica.
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PRIMULACEAE , in See also:

botany, an See also:order of Gamopetalous See also:Dicotyledons belonging to the See also:series Primulales and containing 28 genera with about 350 See also:species. It is See also:cosmopolitan in See also:distribution, but the See also:majority of the species are confined to the temperate and colder parts of the See also:northern hemisphere and many are See also:arctic or alpine. Eight genera are represented in the See also:British See also:flora. The See also:plants are herbs, sometimes See also:annual as in pimpernel (Anagallis arvensis) (fig. 1), but generally perennial as in Primula, (After Wossidlo. From Strasburger's Lehrbuch der Bolanik, by permission of Gustav See also:Fischer.) 1, Flowering See also:branch. 3, See also:Capsule. 2, A See also:flower cut through longi- 4, See also:Seed. tudinally, showing the central 2, 3, 4, Enlarged. See also:placenta. where the plant persists by means of a sympodial rhizome, or in See also:Cyclamen by means of a tuber formed from the swollen hypocotyl. The leaves See also:form a See also:radical rosette as in Primula (See also:primrose, cowslip, &c.), or there is a well-See also:developed aerial See also:stem which is erect, as in species of Lysimachia, or creeping, as in Lysimachia Nummularia (creeping jenny or See also:money-wort). Hottonia (See also:water See also:violet) is a floating water plant with submerged leaves cut into See also:fine linear segments.

The leaves are generally See also:

simple, often with a toothed margin; their arrangement is alternate, opposite or whorled, all three forms occurring in one and the same genus Lysimachia. The See also:flowers are solitary in the See also:leaf-axils as in pimpernel, money-wort, &c., or umbelled as in primrose, where the umbel is sessile, and cowslip, where it is stalked, or in racemes or spikes as in species of Lysimachia. Each flower is subtended by a bract, but there are no bracteoles, and corresponding with the See also:absence of the latter the two first developed sepals stand right and See also:left (fig. 2). The flowers are hermaphrodite and See also:regular with parts in See also:fives (pentamerous) throughout, though exceptions from the pentamerous arrangement occur. The sepals are leafy and persistent; the corolla is generally divided into a longer or shorter See also:tube and a See also:limb which is spreading, as in primrose, or reflexed, as in See also:Diagram of a typical Cyclamen; in Soldanella it is See also:bell-shaped; in sower of Pruuula- Lysimachia the tube is often very See also:short, the ceae• petals appearing almost See also:free; in Glaux the petals are absent. The five stamens See also:spring from the corolla-tube end are opposite to its lobes; this anomalous position is generally explained by assuming that an See also:outer whorl of stamens opposite the sepals has disappeared, though sometimes represented by scales as in Samolus and Soldanella. Another explanation is based on the See also:late See also:appearance of the petals in the floral development and their origin from the backs of the primordia of the stamens; it is then assumed that three alternating whorls only are See also:present, namely, sepals, stamens bearing petal-like dorsal outgrowths, and carpels. The See also:superior ovary—See also:half-inferior in Samolus—bears a simple See also:style ending in a capitate entire stigma, and contains a free-central placenta bearing generally a large number of ovules, which are exceptional in the See also:group Gamopetalae in having two integuments. The See also:fruit is a capsule dehiscing by 5 sometimes ro See also:teeth or valves, or sometimes transversely (a pyxidium) as in Anagallis. See also:Cross See also:pollination is often favoured by dimorphism of the flower, as shown in species of Primula (fig. 3).

The two forms have See also:

long and short styles repectively, the stamens occupying corresponding positions half-way down or at the mouth of the corolla-tube; the long-styled flowers have smaller See also:pollen-grains, which correspond with smaller stigmatic papillae on the short styles. The order is divided into five tribes by characters based on See also:differences in position of the ovules—which are generally semianatropous so that the seed is peltate with the hilum in the centre on one See also:side (or ventral), but sometimes, as in Hottonia and (From Strasburger's Lehrbuch der Botanik.) L, Long-styled flowers. P, Pollen grains, and N, stig- K, Short-styled flowers. matic papillae of long-styled G, Style. form. S, Anthers. p, n, See also:Ditto of short-styled form. (P, N, p, n.) Samolus, anatropous with the hilum basal—together with the method of dehiscence of the capsule and the relative position of the ovary. The See also:chief British genera are Primula, including P. vulgaris, primrose, P. veris, cowslip, P. elatior, oxlip, and the small alpine species P. forinosa, with mealy leaves; Lysimachia, loose strife, including L. Nummularia, money-wort; Anagallis, pimpernel; and Hottonia, water violet.

End of Article: PRIMULACEAE

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