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ORDER I

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Originally appearing in Volume V26, Page 543 of the 1911 Encyclopedia Britannica.
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ORDER I .—CROSSOPTERYGII Paired fins, at least the pectorals, lobate, having an endo-skeletal See also:axis more or less fringed with dermal rays. Mandibular See also:arch suspended from the upper segment of the hyoid arch (hyostylic See also:skull). Splenial See also:bone See also:present. No supraoccipital bone. A pair of large jugular plates, sometimes with small lateral plates and an anterior azygous See also:element, See also:developed in the branchiostegal membrane between the mandibular rami. See also:Heart with a' contractile, multivalvular conus arteriosus; See also:intestine with a See also:spiral See also:valve; See also:air-See also:bladder with pneumatic duct communicating with the ventral See also:side of the See also:oesophagus. Maxillary bone large, toothed, bordering the mouth. Bones of the upper See also:surface of the skull mostly paired. See also:Pectoral arch with both clavicle (so-called infra-clavicle) and cleithrum. Ventral fins inserted far back. With few exceptions (tail of Coelacanthidae, dorsal and caudal fins of Polypteridae) the dermal rays of the unpaired fins more numerous than their endo-skeletal supports, a See also:primitive See also:character also found in the See also:lower Ganoids, but disappearing in the higher. SUB-ORDER I.—OSTEOLEPIDA (Including the Haplistia, Rhipidistia and Actinistia.) Pectoral fins obtusely or acutely lobate, articulating with the pectoral See also:girdle by a single basal endo-skeletal element.

Nostrils on the lower side of the snout. Two dorsal fins. Families: Osteolepidae, Rhizodontidae, Holoptychidae, Coelacanthidae. The scales may be rhombic and thickly coated with ganoine (Osteolepidae) or See also:

cycloid. The vertebral axis is strongly heterocercal in the Osteolepidae and Holoptychidae, and diphycercal or intermediate between the heterocercal and the diphycercal types in the other families; usually acentrous, sometimes with See also:ring-like calcifications (some of the Rhizodontidae). In the Holoptychidae the pectoral fin is extremely similar to that of the Dipneusti of the See also:family Dipteridae, which they resemble closely in See also:form and scaling. Their See also:teeth are remarkable for their complicated structure, resembling that of the Labyrinthodont Batrachians. A pineal foramen is present between the frontal bones in most of the Rhizodontidae. The Osteolepidae were mostly moderate-sized fishes, the largest (Megalichthys) measuring about 4 ft. in length. These Crossopterygians first appear in the Lower Devonian, are abundant in the Upper Devonian, Carboniferous and See also:Permian; in later periods they are represented only by the more specialized Coelacanthidae, which appear in the Lower Carboniferous, and persist as See also:late as the Upper See also:Chalk. SUB-ORDER II.—CLADISTIA Pectoral fin obtusely lobate, with three basal endo-skeletal elements. Nostrils on the upper side of the snout.

A single dorsal fin, formed of a See also:

series of detached rays. A single family: Polypteridae. The existing Crossopterygians which form this sub-order differ very considerably from the See also:extinct Osteolepida, perhaps quite as much as these differ from the Dipneusti. The ventral fins are not lobate, the vertebral See also:column is well ossified and its termination is of the diphycercal type. Spiracles, covered by bony valves, are present on the upper surface of the See also:head. The dorsal fin is unique among fishes, being formed of detached rays consisting of a spine-like fulcra! See also:scale supporting the fringes of the See also:ray; these rays have been regarded, erroneously, as representing so many distinct fins, or " finlets." The scales are bony, rhombic and thickly coated with ganoine. The Polypteridae are confined to tropical See also:Africa and the See also:Nile, and represented by two genera: Polypterus and Calamichthys, the former moderately elongate and provided with ventral fins, the latter serpentiform and devoid of ventrals. We now know ten See also:species of Polypterus, from the Nile, the See also:Congo, the See also:rivers of See also:West Africa, and lakes See also:Chad, See also:Rudolf and See also:Tanganyika, and one of Calamichthys, which inhabits West Africa from the See also:Niger See also:delta to the Chiloango. The largest species of Polypterus reach a length of nearly 4 ft. The See also:young are provided with an See also:external opercular gill very similar to the gills of larval salamanders. The air-bladder acts as an See also:accessory breathing See also:organ, although these fishes are not known ever to leave the See also:water. The development is stated by the late J.

S. Budgett to be even more Batrachian-like than that of the Dipneusti, but the results of the study of the material collected by him shortly before his See also:

death have not yet been published.

End of Article: ORDER I

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