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TORTOISE

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Originally appearing in Volume V27, Page 67 of the 1911 Encyclopedia Britannica.
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TORTOISE . Of the three names generally used for this See also:

order of See also:reptiles, viz. tortoise, turtle and terrapin, the first is derived from the Old See also:French word tortis, i.e. See also:twisted, and was probably applied first to the See also:common See also:European See also:species on See also:account of its curiously See also:bent forelegs. Turtle is believed to be a corruption of the same word, but the origin of the name terrapin is unknown: since the See also:time of the navigators of the 16th See also:century it has been in See also:general use for fresh-See also:water species of the tropics, and especially for those of the New See also:World. The name tortoise is now generally applied to the terrestrial members of this See also:group of animals, and that of turtle to those which live in the See also:sea or pass a See also:great See also:part of their existence in fresh water. They constitute one of the orders of reptiles, the Chelonia: toothless reptiles, with well See also:developed limbs, with a dorsal and a ventral See also:shell composed of numerous bony plates, large firmly fixed quadrates, a See also:longitudinal anal opening and an unpaired copulatory See also:organ. The whole shell consists of the dorsal, more or less See also:convex See also:carapace and the ventral plastron, both portions being joined laterally by the so-called See also:bridge. The carapace is (with the exception of Sphargis) formed by dermal ossifications which are arranged in See also:regular See also:series, viz. a median See also:row (1 nuchal, mostly 8 neurals and 1-3 supracaudal or pygal plates), a right and See also:left row of costal plates which surround and partly replace the ribs, and a consider-able number (about 11 pairs) of marginal plates. The plastron consists of usually 9, rarely II, dermal bones, viz. paired epi-, hyo-, hypo- and xiphi-plastral plates and the unpaired endo-plastral; the latter is homologous with the interclavicle, the epi-plastra with the clavicles, the See also:rest with so-called abdominal ribs of other reptiles. In most Chelonians the bony shell is covered with a hard epidermal coat, which is divided into large See also:shields, commonly called " See also:tortoiseshell." These horny shields or scutes do not correspond in See also:numbers and extent with the underlying bones, although there is a general, vague resemblance in their arrangement; for instance, there is a neural, a paired costal and a paired marginal series. The terminology may be learned from the accompanying illustrations (See also:figs. 1 and 2). The integuments of the See also:head, See also:neck, tail and limbs are either soft and smooth or scaly or tubercular, frequently with small osseous nuclei.

All the bones of the See also:

skull are suturally See also:united. The dentary portion of the mandible consists of one piece only, both halves being completely fused together. s The See also:pectoral See also:arch remains See also:separate in the median See also:line; it consists of the coracoids, which slope backwards, and the scapulae, which stand upright and often abut against the inside of the first pair of costal plates. Near the glenoid cavity for the humerus arises from the scapula a See also:long See also:process which is directed transversely towards its See also:fellow; it represents the acromial process of other vertebrates, although so much enlarged, and is neither the precoracoid, nor the clavicle, as stated by the thought-less. The tail is still best developed in the Chelydridae, shortest in the Trionychoidea. Since it contains the large copulatory organ, it is less reduced in the See also:males. No Chelonians possess the slightest traces of See also:teeth, but their jaws are provided with horny sheaths, with hard and See also:sharp edges, forming a See also:beak. The number of Chelonians known at See also:present may be estimated at about 200, the fresh-water species being far the most numerous, and are abundant in well-watered districts of the tropical and sub-tropical zones. Their number and variety decrease beyond the tropics, and in the See also:north they disappear entirely about the 5oth parallel in the western and about the 56th in the eastern hemisphere, whilst in the See also:southern hemisphere the terrestrial forms seem to advance to 36° S. only. The marine turtles, which are spread over the whole of the See also:equatorial and sub-tropical seas, sometimes stray beyond those limits.

End of Article: TORTOISE

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