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CONVOLVULACEAE

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Originally appearing in Volume V07, Page 67 of the 1911 Encyclopedia Britannica.
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CONVOLVULACEAE , a botanical natural See also:

order belonging to the See also:series Tubiflorae of the sympetalous See also:group of See also:Dicotyledons. It contains about 40 genera with more than See also:i000 See also:species, and is found in all parts of the See also:world exdept the coldest, but is especially well See also:developed in tropical See also:Asia and tropical See also:America. The most characteristic members of the order are See also:twining See also:plants with generally smooth See also:heart-shaped leaves and large showy See also:white or See also:purple See also:flowers, as, for instance, the greater bindweed of See also:English hedges, Calystegia se See also:plum, and many species of the genus I pomaea, the largest of the order, including the " convolvulus See also:major " of gardens, and See also:morning See also:glory. The creeping or trailing type is a See also:common one, as in the English bindweed (Convolvulus arvensis), which has also a tendency to climb, and Calystegia Soldanella, the See also:sea-bindweed, the See also:long creeping See also:stem of which forms a See also:sand-binder on English seashores; a widespread and efficient tropical sand-binder is Ipomaea Pes-Caprae. One of the commonest tropical weeds, Evolvulus alsinoides, has slender, long-trailing stems with small leaves and flowers. In hot dry districts such as See also:Arabia and See also:north-See also:east tropical See also:Africa, genera have been developed with a See also:low, much-branched, dense, shrubby See also:habit, with small hairy leaves and very small flowers. An exceptional type in the order is represented by Humbertia, a native of See also:Madagascar, which forms a large See also:tree. The See also:dodder (q.v.) is a genus (Cuscuta) of leafless parasites with slender See also:thread-like twining stems. The flowers stand singly in the See also:leaf-axils or See also:form few or many flowered cymose inflorescences; the flowers are sometimes crowded into small heads. The bracts are usually See also:scale-like, but sometimes foliaceous, as for instance in Calystegia, where they are large and envelop the calyx. The parts of the See also:flower are in See also:fives in calyx, corolla and stamens, followed by two carpels which unite to form a See also:superior ovary. The sepals, which are generally See also:free, show much variation in See also:size, shape and covering, and afford valuable characters for the distinction of genera or sub-genera.

The corolla is generally See also:

funnel-shaped, more rarely See also:bell-shaped or tubular; the See also:outer See also:face is often marked out in See also:longitudinal areas, five well-defined areas tapering from See also:base to See also:apex, and marked with longitudinal striae corresponding to the See also:middle of the petals, and alternating with five non-striated weaker triangular areas; in the bud the latter are folded inwards, the stronger areas being exposed and showing a twist to the right. The slender filaments of the stamens varywidely, often in the same flower; the anthers are linear to ovate in shape, attached at the back to the filament, and open lengthwise. Some importance attaches to the form of the See also:pollen grains; the two See also:principal forms are ellipsoidal with longitudinal bands forming the Convolvulus-type, and a spherical form with a spiny See also:surface known as the Ipomaea-type. The ovary is generally two-chambered, with two inverted ovules See also:standing See also:side by side at the inner See also:angle of each chamber. The See also:style is See also:simple or branched, and the stigma is linear, capitate or globose in form; these See also:variations afford means for distinguishing the different genera. The See also:fruit is usually a See also:capsule opening by valves; the seeds, where four are developed, are each shaped like the quadrant of a See also:sphere; the See also:seed-coat is smooth, or sometimes warty or hairy; the embryo is large with generally broad, folded, notched or bilobed cotyledons surrounded by a horny endosperm. Cuscuta has a thread-like, spirally See also:twisted embryo with no trace of cotyledons. The large showy flowers are visited by See also:insects for the See also:honey which is secreted by a See also:ring-like disk below the ovary; large- Convolvulus sepium, slightly reduced. 1. Flower cut vertically. 4. Embryo taken out of seed.

2. Fruit, slightly reduced. 5. See also:

Horizontal See also:plan of arrange- 3. Seed cut lengthwise showing ment of flower. embryo. flowered species of Ipomaea with narrow tubes are adapted for the visits of honey-seeking birds. The largest genus, Ipomaea, has about 400 species distributed throughout the warmer parts of the See also:earth. Convolvulus has about 150 to 200 species, mainly in temperate climates; the genus is principally developed in the Mediterranean See also:area and western Asia. Cuscuta contains nearly roo species in the warmer and temperate regions; two are native in See also:Britain. The tubers of Ipomaea See also:Bat at as are See also:rich in See also:starch and See also:sugar, and, as the " sweet See also:potato," form one of the most widely distributed foods in the warmer parts of the earth. Several members of the order are used medicinally for the strong purging properties of the milky juice (latex) which they contain; See also:scammony is the dried latex from the underground stem of Convolvulus Scammonia, a native of the See also:Levant, while See also:jalap is the product of the tubercles of Exogonium Purga, a native of See also:Mexico. Species of Ipomaea (morning glory), Convolvulus and Calystegia are cultivated as ornamental plants.

Convolvulus arvensis (bindweed) is a pest in See also:

fields and gardens on See also:account of its wide-spreading underground stem, and many of the dodders (Cuscuta) cause damage to crops.

End of Article: CONVOLVULACEAE

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