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AQUARIUM (plural aquaria)

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Originally appearing in Volume V02, Page 239 of the 1911 Encyclopedia Britannica.
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AQUARIUM (plural aquaria) , the name given to a receptacle for a marine See also:flora and See also:fauna. Until comparatively recently, AQUARIUM 237 aquaria were little more than domestic toys, or show-places of a popular See also:character, but they have now not only assumed a profound scientific importance for the convenient study of anatomical and physiological problems in marine See also:botany and See also:zoology, but have also attained an economic value, as offering the, best opportunities for that study of the habits and environment of marketable See also:food-See also:fish without which no steps for the improvement of See also:sea-See also:fisheries can be safely taken. The numerous " zoological stations " which have sprung up, chiefly in See also:Europe and the See also:United States, but also in the See also:British colonies and See also:Japan, often endeavour to unite these two aims, and have in many cases become centres of experimental See also:work in problems See also:relating to fisheries, as well as in less directly See also:practical subjects. Of these stations, the See also:oldest and the most important is that at See also:Naples, which, though designed for purely scientific See also:objects, also encourages popular study by means of a public aquarium. The following See also:account (1902) of this station by Dr W. Giesbrecht, a member of the See also:staff, will serve to show the methods and aims, and the complex and expensive equipment, of a See also:modern aquarium: " The zoological station at Naples is an institution for the See also:advancement of biological science—that is, of See also:comparative See also:anatomy, zoology, botany, See also:physiology. It serves this end by providing the biologist with the various objects of his study and the necessary appliances; it is not a teaching institution. The station was founded by Dr Anton Dohrn, and opened in the See also:spring of 1874; it is the oldest and largest of all biological stations, of which there are now about See also:thirty in existence. Its two buildings See also:ate situated near the seashore in the western See also:town See also:park (See also:Villa Nazionale) of Naples. The older and larger one, 33 metres See also:long, 24 M. deep, 16 m. high, contains on the ground See also:floor the aquarium, which is open to the public. On the first floor there is, facing See also:south, the See also:principal library, ornamented with See also:fresco paintings, and, facing See also:north, a large See also:hall containing twelve working tables, several smaller rooms and the secretarial offices. On the second floor is the physiological laboratory, and on the third floor the small library, a hall with several working tables, and the dark rooms used in developing photographs.

The ground floor of the smaller See also:

building, which was finished in 1887, contains the rooms in which the animals are delivered", sorted and pre-served, and the fishing tackle kept, together with the workshop of the engineer; on the first and second floors are workrooms, amongst others the botanical laboratory; on the third floor are See also:store-rooms. In the See also:basement of both buildings, which is continued underneath the See also:court, there are sea-See also:water cisterns and filters, engines and store-rooms. The materials for study which the station offers to the biologist are specimens of marine animals and See also:plants which abound in the western See also:part of the Mediterranean, and especially in the Gulf of Naples. To obtain these, two See also:screw-steamers and several See also:rowing boats are required, which are moored in the See also:harbour of Mergellina, situated See also:close by. The larger steamer, ` Johannes See also:Muller' (15 M. long, 21 M. wide, 1 m. See also:draught), which can See also:steam eight to ten See also:English See also:miles per See also:hour, is provided with a steam dredge working to a See also:depth of eighty fathoms. From the small steamer, ` See also:Frank See also:Balfour,' and the rowing boats, the fishing is done by means of See also:tow-nets. Besides these there are fishermen and others who daily See also:supply living material for study. The See also:plankton (small floating animals) is distributed in the See also:morning, other animals as required. The animals brought in by the fishermen are at once distributed amongst the biologists, whereas the material brought up by the dredges is placed in See also:flat revolving wooden vessels, so as to give the smaller animals See also:time to come out of their hiding-places. The students who work in the station have the first claim on specimens of plants and animals; but specimens are also sup-plied to museums, laboratories and See also:schools, and to individuals engaged in See also:original See also:research elsewhere. Up to the See also:present time about 4000 such parcels have been despatched, and not infrequently live specimens of animals are sent to distant places. This See also:side of the work has been of very See also:great value to See also:science.

The principal appliances for study with which the station provides the biologist are workrooms furnished with the apparatus and chemicals necessary for anatomical research and physiological experiments and tanks. Every student receives a tank for his own See also:

special use. The large tanks of the principal aquarium are also at his disposal for purposes of observation and experiment if necessary. "The water in the tanks is kept fresh by continual circulation, and is thus charged with the See also:oxygen necessary to the See also:life of the organisms. It is not pumped into the tanks directly from the sea, but from three large cisterns (containing 300 cubic metres), to which it again returns from the tanks. The water wasted or evaporated during this See also:process is replaced by new water pumped into the cisterns directly from the sea. The water flows from the large cisterns into a smaller cistern, from which it is distributed by means of an electric See also:pump through vulcanite or See also:lead pipes to the various tanks. The water with which the tanks on the upper floors are filled is first pumped into large wooden tanks placed beneath the roof, thence it flows, under almost See also:constant pressure, into the tanks. The water circulated in this manner contains by far the largest number of such animals as are capable of living in captivity in See also:good See also:condition. Some of them even increase at an undesirable See also:rate, and it some-times happens that See also:young Mytilus or Ciona stop up the pipes; in laying these, therefore, due regard must be had to the arrangements for cleaning. For the cultivation of very delicate animals it is necessary to keep the water absolutely See also:free from harmful bacteria; for this purpose large See also:sand-filters have lately been. placed in the See also:system, through which the water passes after leaving the cisterns. Each of the smaller cisterns, which are fixed in the workrooms, consist of two water-tanks, placed one above the other; their frames are of wrought See also:iron and the walls generally of See also:glass.

Vessels containing See also:

minute animals can be placed between these two tanks, receiving their water through a See also:siphon from the upper tank; the water afterwards flows away into the See also:lower tank. " The twenty-six tanks of the public aquarium (the largest of which contains 112 cubic metres of water) have See also:stone walls, the front portion alone being made of glass. As the tanks hold a very large number of animals in proportion to the quantity of water, they require to be well aerated. The pipes through which the water is conducted are therefore placed above the See also:surface of the water, and the fresh supply is driven through them under strong pressure. A large quantity of See also:air in the See also:form of See also:fine bubbles is thus taken to the bottom of the tank and distributed through the entire See also:mass of water. Should the organisms which it is desired to keep alive be very minute, there is a danger of their being washed away by the circulating water. To obviate this, either the water which flows away is passed through a strainer, or the water is not changed at all, air being driven through it by means of an apparatus put into See also:motion by the drinking-water supply. " The library contains about 9000 volumes, which students use with the help of a slip See also:catalogue, arranged according to authors. The station has published at intervals since 1879 two See also:periodicals treating of the organisms of the Mediterranean. One is Fauna and Flora See also:des Golfes von Neapel, the other Mittheilungen aus der zoologischen Station zu Neapel. The former consists of mono-graphs in which special See also:groups of animals and plants are most exhaustively treated and the Mediterranean See also:species portrayed according to life in natural See also:colours; up to the present time twenty-one zoological and five botanical monographs have appeared, making altogether 1200 4to sheets with about 400 plates. Of the 111ittlieilungen, which contain smaller articles on organisms of the Mediterranean, fourteen volumes in 8vo have been published.

The station also publishes a Zoologischer Jahresbericht, which at first treated of the entire See also:

field of zoology, but since 1886 has been confined principally to comparative anatomy and ontogeny; it appears eight to nine months after the end of the See also:year reported. The See also:Guide to the Aquarium, with its descriptions and numerous pictures, is meant to give the See also:lay visitor an See also:idea of the marine See also:animal See also:world. "There are about See also:forty officials, amongst them six zoologists, one physiologist, one secretary, two draughtsmen, one engineer. The station is a private institution, open to biologists of all nations under the following conditions: there are agreements with the governments of See also:Austria, See also:Baden, See also:Bavaria, See also:Belgium, See also:Hamburg, See also:Holland, See also:Hesse, See also:Italy, See also:Prussia, See also:Russia, See also:Saxony, See also:Switzerland, See also:Hungary, See also:Wurttemberg, the See also:province of Naples, and the See also:universities of See also:Cambridge, See also:Oxford, See also:Strassburg, See also:Columbia See also:College (New See also:York), and the British Association for the Advancement of Science, the Smithsonian Institution, and a society of See also:women in the United States of North See also:America (formerly also with See also:Bulgaria, See also:Rumania, See also:Spain, the See also:Academy of Sciences in See also:Berlin, See also:Williams College, University of See also:Pennsylvania), by virtue of which the governments and corporate bodies named have the right, on See also:payment of £loo per annum, to send a worker to the station; this places at his disposal a ` table' or workplace, furnished with all the necessary appliances and materials as set down in the agreement. At present there are agreements for thirty-three tables, and since the See also:foundation of the station nearly 1200 biologists have worked there. The current expenses are paid out of the table-rents, the entrance fees to the public aquarium, and an See also:annual subvention paid by the See also:German See also:empire." In See also:England a station on similar lines, but on a smaller See also:scale, is maintained at See also:Plymouth by the Marine Biological Association of the United See also:Kingdom, with the help of subsidies from the See also:government and the Fishmongers' See also:Company. Little difficulty is experienced in maintaining, breeding and rearing fresh-water animals in captivity, but for many various reasons it is only by unremitting See also:attention and foresight that most marine animals can be kept even alive in aquaria, and very few indeed can be maintained in a condition healthy enough to breed. Much experience, however, has been gained of See also:late years at considerable expense, both in England and abroad. In starting a marine aquarium of whatever See also:size, it should be obvious that the first See also:consideration must be a supply of the purest possible water, as free as may be, not only from See also:land-drainage and sewage, but also from such suspended matters as See also:chalk, fine sand or mud. This is most ideally and economically secured by placing the station a few feet above high-water See also:mark, in as sheltered a position as possible, on a rocky See also:coast, pumping from the sea to a large See also:reservoir above the station, and allowing the water to circulate gently thence through the tanks by gravity (Banyuls). At an inland aquarium (Berlin, Hamburg), given pure water in the first instance, excellent if less See also:complete results may nevertheless. be obtained. The next consideration is the method by which oxygen is to be supplied to the organisms in the aquarium.

Of the two methods hitherto in use, that of pumping a See also:

jet of air into tanks otherwise stagnant or nearly so (See also:Brighton), while supplying sufficient oxygen, has so many other disadvantages, that it has not been employed regularly in any of the more modem aquaria. It is, however, still useful in aerating quite small bodies of water in which See also:hardy and minute organisms can be isolated and kept under See also:control. In the other method, now in See also:general use, a fine jet of water under pressure falls on to the surface of the tank; this carries down with it a more than sufficient air-supply, See also:analysis showing in some cases a higher percentage of oxygen in aquarium water than in the open sea. The water supply is best effected by gravity from reservoirs placed above the tanks, but may be also achieved by See also:direct pumping from See also:low reservoirs or from the sea to the tanks. Provided that an unlimited supply of pure water can be obtained cheaply, the overflow from the tanks is best run to See also:waste; but in aquaria less fortunately placed, it returns to a storage low-level reservoir, from which it is again pumped, thus circulating See also:round and round (Naples, Plymouth). The storage reservoirs should be in all cases very large in comparison with the bulk of water in circulation; if practicable, they should be excavated in See also:rock, and lined with the best See also:cement. There is no See also:reason why they should not be shallow, exposed to See also:light and air, and cultivated as rock-pools by the introduction of seaweeds and small animals, but they must then be screened from See also:rain,. See also:cold and dust. The pumps used in circulation will be less likely to kill minute animals if of the plunger or See also:ram type, rather than rotary, and should be of See also:gun-See also:metal- or one of the new See also:bronze-See also:alloys which take a See also:patina in See also:salt water. For the circulating pipes many materials have been tried. Vulcanite is not only expensive and brittle, but has other disadvantages; See also:common iron pies, coated internally with cement or See also:asphalt or glazed internally, with all unions and See also:joints cemented, have been used with more or less success. Probably best of all is common lead piping, the joints being served with red-lead; water should be circulated through such pipes till they become coated with in-soluble carbonate, for some time before animals are put into the tanks. For small installations glass may be used, the joints being made with marine See also:glue or other suitable cement.

In building the tanks themselves, regard must be had to their special purposes. If intended for show-tanks for popular admiration, or for the study of large animals, they must be large with a See also:

plate-glass front; for See also:ordinary scientific work small tanks with all sides opaque are preferable from every point of view. According to their character, size and position, fixed tanks may be of See also:brickwork, See also:masonry or rock, coated in each See also:case with cement; asphalting the sides offers no particular advantages, and often gives rise to great trouble and expense. All materials, and especially the cements, must be of the finest quality procurable. For smaller and movable tanks, See also:slate slabs bolted or screwed together have some disadvantages, notably those of expense, See also:weight and brittleness, but are often used. Better, cheaper and lighter, if less permanent, are tanks of See also:wood bolted together, pitched internally. Glass See also:bell-jars, useful in particular cases, should generally have their sides darkened, except when required for observation. See also:Provision should always be made for cleaning every part of the tanks, pipes and reservoirs; all rock-work in tanks should therefore be removable. As regards the See also:lighting of fixed tanks, it should always be directly from above. In all tanks with glass sides, whether large or small, as much light as possible should be kept from entering through the glass; otherwise, with a side-light, many animals become restless, and See also:wear themselves out against the glass, affected by even so little light as comes through an opposite tank. In cases where distance from the sea or other causes make it impracticable to allow the overflow from the tanks to run to waste, special precautions must be taken to keep the water pure. Chemically speaking, the See also:chief character of the water in an aquarium circulation, when compared with that of the open sea, lies in the excessive quantity of See also:nitrogen present in various forms, and the reduced alkalinity; these two being probably connected.

The excess of nitrogen is referable to dead animals, to waste food and to the excreta of the living organisms. The first two of these See also:

sources of contamination may be reduced by care and cleanliness, and by the See also:maintenance of a flow of water sufficient to prevent the excessive See also:accumulation of sediment in the tanks. The following experiment shows the rapid rise of nitrogen if unchecked. A tank with a considerable fauna was isolated from the general circulation and aerated by four air-jets, except during See also:hours 124—166 bf the experiment; See also:column I. shows per See also:Ioo,000 the nitrogen estimated as See also:ammonia, column II. the See also:total inorganic nitrogen: I. II. Sea-water at source of original 0.003 supply . 0.001 Aquarium water in tank at com- 0.400 mencement of experiment . 0.012 After 221 hours . 0.020 . 11 75 0.025 I.200 11 93 „ 0.019 . 1211 „ 0.012 11 141 „ . 0.015 2.200 165 0.025 ..

169 „ . 0.025 11 189 . .0.012 During this time the alkalinity was reduced to the See also:

equivalent of 30 mg. CaCO3 per litre, ocean water having an alkalinity equivalent to 50-55 mg. per litre. It has been suggested that the organic nitrogen becomes oxidized into nitrous, then into nitric See also:acid, which lowers the carbonate values. A great See also:deal of reduction of this nitrogenous contamination can be effected by filtration, See also:AQUAVIVA 239 a method first introduced successfully at Hamburg, where a most thriving aquarium has been maintained by the See also:local Zoological Society for many years on the circulation principle, new water being added only to compensate for waste and evaporation. The filters consist of open See also:double boxes, the inner having a bottom of perforated slate on which rests rough See also:gravel; on the latter is fine gravel, then coarse, and finally fine sand. Filtration may be either upwards or downwards through the inner See also:box to the See also:outer. Such filters, intercalated between tanks and reservoir, have been shown by analysis to stop a very large proportion of nitrogenous See also:matter. It is doubtful whether aquarium water will not always show an excess of nitrogenous compounds, but they must be kept down in every way possible. In small tanks, well lighted, seaweeds can be got to flourish in a way that has not been found practicable in large tanks with a circulation; these, with Lamellibranchs and small See also:Crustacea as scavengers, will be found useful in this connexion. Slight or occasional circulation should be employed here also, to remove the film of dust and other matters, which otherwise covers the surface of the water and prevents due oxygenation.

In such small tanks for domestic use the fauna must be practically limited to bottom-living animals, but for purposes of research it is often desired to keep alive larval and other surface-See also:

swimming animals (plankton). In this case a further difficulty is presented, that of helping to suspend the animals in the water, and thus to avoid the exhaustion and See also:death which soon follow their unaided efforts to keep off the bottom; this See also:duty is effected in nature by specific gravity,, See also:tide and surface current. In See also:order to deal with this difficulty a See also:simple but efficient apparatus has been devised by Mr E. T. See also:Browne; a " plunger," generally a glass plate or See also:filter See also:funnel, moves slowly up and down in a bell-See also:jar or other small tank, with a See also:period of See also:rest between each stroke; the See also:motive See also:power is obtained through a simple bucketand-siphon arrangement worked by the overflow from other tanks. This apparatus (first used at the Plymouth Laboratory of the Marine Biological Association in 1897, and since introduced into similar institutions), by causing slight eddies in the water, keeps the floating fauna in suspension, and has proved very successful in rearing larvae and in similar work. (G. H.

End of Article: AQUARIUM (plural aquaria)

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